Gobio gobio

  • Scientific name
  • Gobio gobio (Linnaeus, 1758)

  • Common name
  • Gudgeon

  • Family
  • Cyprinidae

  • External links
  • Fishbase
Trait completeness 96%
Total data222
References36
Image of Gobio gobio

Author: Fabrice Téletchéa
License: All rights reserved

Traits detail



Egg (100%)


Trait id Trait Primary data Secondary Data References
4 Egg adhesiveness Adhesive Adhesive Spillmann, 1961
4 Egg adhesiveness Adhesive [Eggs adhere to pebbles, weeds, tree roots and sodden leaves] Adhesive Kennedy and Fitzmaurice, 1972
4 Egg adhesiveness Adhesive Adhesive Bruslé and Quignard, 2001
4 Egg adhesiveness Slightly adhesive Adhesive Rinchard, 1996
4 Egg adhesiveness Adhesive Adhesive Billard, 1997
4 Egg adhesiveness Highly adhesive Adhesive Rossechi and Kestemont, 2001
4 Egg adhesiveness Sticky Adhesive Penaz and Prokes, 1978
4 Egg adhesiveness Fish eggs were unsticked enzymatically using alcalase for three minutes Adhesive Palikova and Krejci, 2006
4 Egg adhesiveness Les œufs pondus adhérent aux graviers Non-Adhesive Brunet and Hoestlandt, 1972
5 Incubation time 14-28 21.0 days Spillmann, 1961
5 Incubation time 5-6 at 20°C 5.5 days Bruslé and Quignard, 2001
5 Incubation time 7-8 days at 17.5°C 7.5 days Kennedy and Fitzmaurice, 1972
5 Incubation time In the Mohelno locality, it lasted 7 days at daily mean water temperature of 16.9°C and in laboratory, 150 hours (5days) at 19.5°C. At mean water temperature of 20.14°C, the first embryos hatched in the 190th hour of development, i.e. on the 5th days after insemination of the eggs. Hatcing ended in the 164th hour of development, i.e., on the 7th day after insemination of the eggs. Other authors: 8.5 daysat 16°C, 8 days at 17.5°Cand 6 at 18°C 7.0 days Penaz and Prokes, 1978
5 Incubation time The hatching of eggs from females not treated hormonally started on average 71 hours after fertilization with the range of 66 to 79 hours. Most eggs hatched within 39 hours from the start of hatching. All embryos hatched within 136 hours after fertilization. 71.0 days Palikova and Krejci, 2006
5 Incubation time 6 days at 20°C 6.0 days Brunet and Hoestlandt, 1972
7 Degree-days for incubation 125 125.0 °C * day Rinchard, 1996
7 Degree-days for incubation 125 125.0 °C * day Bruslé and Quignard, 2001
7 Degree-days for incubation 120-140 [7-8 days at 17.5°C] 130.0 °C * day Kennedy and Fitzmaurice, 1972
7 Degree-days for incubation The mean duration of incubation was 133.4 DD at 20.4°C, also described at 6 days at 20°C (125 DD) 133.4 °C * day Penaz and Prokes, 1978
7 Degree-days for incubation 125°D 125.0 °C * day Brunet and Hoestlandt, 1972
6 Temperature for incubation 17-18 17.5 °C Kennedy and Fitzmaurice, 1972
6 Temperature for incubation 20°C 20.0 °C Bruslé and Quignard, 2001
6 Temperature for incubation Artificially inseminated eggs were incubated on Petri dishes at water temperatures varying between 18.1 and 20.5°C 18.1 °C Penaz and Prokes, 1978
6 Temperature for incubation Fish eggs were incubated at 21 ± 0.3°C 21.0 °C Palikova and Krejci, 2006
6 Temperature for incubation 19-20°C 19.5 °C Brunet and Hoestlandt, 1972
2 Egg size after water-hardening Mainly 1.3, varying between 1.3-1.35 [Drifting eggs] 1.325 mm Copp, 2002b
2 Egg size after water-hardening At water temperature of 19°C, the water uptake lasted about 45 minutes and the eggs increased their diameter to 1.29 mm 1.29 mm Penaz and Prokes, 1978
2 Egg size after water-hardening 1.5 [Not specified] 1.5 mm Kamler and Wolnicki, 2006
2 Egg size after water-hardening Fertilized fish eggs in a swollen stated had 1.51 mm (1.42-1.62) in diameter and their mass was 1.48 mg on average (1.22-1.87) 1.52 mm Palikova and Krejci, 2006
2 Egg size after water-hardening Les œufs sont blancs tanslucides et mesurent 1.4 à 1.8 mm de diamètre (moyenne 1.5 mm), ceci rejoint les observations en Irlande où les œufs de goujons mesurent de 1.30 à 1.65 mm 1.4 mm Brunet and Hoestlandt, 1972
3 Egg Buoyancy Demersal Demersal Kennedy and Fitzmaurice, 1972
3 Egg Buoyancy Demersal Demersal Tyler and Sumpter, 1996
1 Oocyte diameter 1.375-1.625 [Ova] 1.5 mm Kennedy and Fitzmaurice, 1972
1 Oocyte diameter 1.5-2 1.75 mm Bruslé and Quignard, 2001
1 Oocyte diameter 2 [Not specified] 2.0 mm Spillmann, 1961
1 Oocyte diameter 1.2 1.2 mm Rossechi and Kestemont, 2001
1 Oocyte diameter 1.4-1.8 1.6 mm Fishbase, 2006
1 Oocyte diameter 1.126 [Mean egg size, not specified] 1.126 mm Rosecchi, 2001
1 Oocyte diameter 1.2 1.2 mm Tyler and Sumpter, 1996
1 Oocyte diameter 0.73-1.76, up to 2.20 ? [Average diameter of the largest oocyte in fully developed ovaries] 1.245 mm Vila-Gispert and Moreno-Amich, 2002
1 Oocyte diameter 1.8 [Not specified] 1.8 mm Copp, 2002b
1 Oocyte diameter Non-inseminated, freshly stripped gudgeon eggs are 0.99 mm in diameter 0.99 mm Penaz and Prokes, 1978
1 Oocyte diameter Unfertilized and not swollen fish eggs measured 1.24 mm on average, with the mean range from 1.17 to 1.32 mm (and their mass averaged 0.70 mg, 0.67 to 0.78) 1.24 mm Palikova and Krejci, 2006

Larvae (100%)


Trait id Trait Primary Data Secondary Data References
11 Temperature during larval development 20-25 [At 28 optimal growth but problem with anoxia and pathology] 22.5 °C Chemillier, 1995
11 Temperature during larval development 18-20 19.0 °C Kennedy and Fitzmaurice, 1972
11 Temperature during larval development Reared at 20°C 20.0 °C Wolnicki, 2005
10 Reaction to light Benthic larvae are photophobic Photophobic Mann, 1996
12 Sibling intracohort cannibalism Not any cannibalism has ever been observed Absent Chemillier, 1995
13 Full yolk-sac resorption 90-100 [5 days at 18-20°C] 95.0 °C * day Kennedy and Fitzmaurice, 1972
13 Full yolk-sac resorption La vésicule est résorbée en quatre jours (à + 20°C) et les alevins de notre élevage mesurent 5.15 (5 à 5.5 mm) alors que l'alevin d'Irlande atteint 6 mm 20.0 °C * day Brunet and Hoestlandt, 1972
14 Onset of exogeneous feeding 60 [3 days at 18-20°C] 19.0 °C * day Kennedy and Fitzmaurice, 1972
8 Initial larval size 5.45 5.45 mm Kennedy and Fitzmaurice, 1972
8 Initial larval size 4-5 4.5 mm Bruslé and Quignard, 2001
8 Initial larval size 4.5-5.5 5.0 mm Rinchard, 1996
8 Initial larval size 4.7 4.7 mm Chemillier, 1995
8 Initial larval size 3.0-4.2 3.6 mm Penaz and Prokes, 1978
8 Initial larval size Mean of 4.3 (4 to 4.7 mm). […] In Ireland, length of 4.68 to 5.45 mm 4.3 mm Brunet and Hoestlandt, 1972
9 Larvae behaviour Tend to keep to the bottom. where they rest upright with the spread out of their pectoral fins Demersal Kennedy and Fitzmaurice, 1972
9 Larvae behaviour Benthic larvae Demersal Mann, 1996
9 Larvae behaviour The hatched embryos mostly lie still on the bottom Demersal Penaz and Prokes, 1978

Female (100%)


Trait id Trait Primary Data Secondary Data References
18 Female sexual dimorphism Tubercles are present mostly on the head, and in the largest individuals also on scales of back Present Witkowski and Rogowska, 1991
24 Maximum GSI value 16.53 ± 1.53 [Idem as Spawning value, May-June] 16.53 percent Rinchard, 1993
24 Maximum GSI value 23-28.6 25.8 percent Bruslé and Quignard, 2001
24 Maximum GSI value 16.5% [June, and July] 16.5 percent Kestemont, 1987
24 Maximum GSI value 17.1 [April, May] 17.1 percent Rosecchi, 2001
24 Maximum GSI value Huge variations between females, from 4 to 22, most between 12-18% in May 15.0 percent Mann, 1980
24 Maximum GSI value 13.5 [End of May in the control group] 13.5 percent Kestemont, 1990
25 Oogenesis duration 9-10 [October to June] 9.5 months Kestemont, 1987
19 Relative fecundity 180-849 514.5 thousand eggs/kg Bruslé and Quignard, 2001
19 Relative fecundity 200-250: 500-4500 [Age 2, weight 10g], 1500-9000 [Age 3, 20-25 g] 225.0 thousand eggs/kg Kestemont and Mélard, 1994
27 Age at sexual maturity 2-3 [Both sex] 2.5 years Kennedy and Fitzmaurice, 1972
27 Age at sexual maturity 2 2.0 years Bruslé and Quignard, 2001
27 Age at sexual maturity 2 [Sometimes 1, rarely 3-4, Box sex] 3.5 years Rosechhi and Kestemont, 2001
27 Age at sexual maturity 2-3 [male] 2.5 years Fishbase, 2006
27 Age at sexual maturity 1 [Bot sex] 1.0 years Rosecchi, 2001
27 Age at sexual maturity 3-4 [Males] 3.5 years Environment agency, 1996
27 Age at sexual maturity Nearly all fish were mature at the end of their third year, and approximatively 64% of all males after two years 2.0 years Mann, 1980
26 Resting period 1-2 [August-September] 1.5 months Kestemont, 1987
26 Resting period 2-3 [From July through September] 2.5 months Rinchard, 1993
26 Resting period From July, the ovary started a recovery phase and only contained stage 1 and 2 ooctyes. One ot two months of quiscence following summer spawning 1.0 months Kestemont, 1990
22 Onset of oogenesis Mid-October [Greatly decreased between November to April] ['April', 'October', 'November'] Kestemont, 1987
22 Onset of oogenesis October to March slight increase from 3 to 4.59 ['October', 'March'] Rinchard, 1993
22 Onset of oogenesis Increase in October ['October'] Mann, 1980
22 Onset of oogenesis In October, all fishes were at the same stage of development and only contained stage 1 and 2 ooctyes. Stage 3, reached in November, continued until April as the most advanced stage. ['April', 'October', 'November'] Kestemont, 1990
23 Intensifying oogenesis activity May ['May'] Kestemont, 1987
23 Intensifying oogenesis activity April-May ['April', 'May'] Rinchard, 1993
23 Intensifying oogenesis activity April [From 6.3 in March to 17.6 in April] ['April', 'March'] Rosecchi, 2001
23 Intensifying oogenesis activity April ['April'] Mann, 1980
23 Intensifying oogenesis activity Enlargement of ooctyes by accumulation of yolk globules caused a marked increase in GSI in May ['May'] Kestemont, 1990
21 Oocyte development Between group-synchronous and synchronous Group-synchronous Rinchard, 1996
21 Oocyte development Asynchronous Asynchronous Bruslé and Quignard, 2001
21 Oocyte development Asynchonous Asynchronous Kestemont, 1987
21 Oocyte development Develops oocyes asynchronously Asynchronous Kestemont, 1990
21 Oocyte development Développement asynchrone Asynchronous Beelen, 1998/1999
20 Absolute fecundity 1-3 2.0 thousand eggs Spillmann, 1961
20 Absolute fecundity 0.5-15 7.75 thousand eggs Rinchard, 1996
20 Absolute fecundity 4.8-20.8 [Great Britain] and 0.5-3 [France] 12.8 thousand eggs Bruslé and Quignard, 2001
20 Absolute fecundity 1.987-3.240 for three populations up to 15.950 in England [Average number of vitellogenic oocyes of mature females in a single spawning season] 2.6135 thousand eggs Vila-Gispert and Moreno-Amich, 2002
20 Absolute fecundity 2.5-6.5 4.5 thousand eggs Environment agency, 1996
20 Absolute fecundity Mean number of eggs per fish in successive age groups: 4812 [Age 2], 9787 [Age 3], 15950 {Age 4], 20823 [Age 5] in Frome, 5044 [Age 3], 4472 [Age 4], and 6781 [Age 5] in Thames, 500 [Age 2], 1500 [Age 3], 3000 [Age 4] and 4-5000 [Age 6] in Nivelle, 1721 [Age 3], 2053 [Age 4], 2886 [Age 5], 3585 [Age 6], 5610 [Age 7] in Dunajec 2502.0 thousand eggs Mann, 1980
20 Absolute fecundity Mean of 4 643, range 629-14 600 321.5 thousand eggs Banbura and Koszalinski, 1991
20 Absolute fecundity Categorized as between 2000 and 100000 eggs per reproductive cycle 2000.0 thousand eggs Cattanéo, 2001
20 Absolute fecundity 500-4500 [Age 2], 1500-9000 [Age 3], 3000-15000 [Age 4] 2500.0 thousand eggs Kestemont and Mélard, 1994
20 Absolute fecundity On peut admettre que la femelle agée de 1+ pond 500 œufs, celle de 2+ 15000, celle de 3+ 3000 et celle de 5+ 4000 à 5000 1.0 thousand eggs Brunet and Hoestlandt, 1972
17 Weight at sexual maturity Gudgeon generally reaches a size of 8-10 cm and a weight of 10 g at 2 years, 12-13 cm and 20-25 g at 3 years 9.0 kg Kestemont and Mélard, 1994
16 Length at sexual maturity 7.9 is the smallest mature female 7.9 cm Kennedy and Fitzmaurice, 1972
16 Length at sexual maturity 8-10 9.0 cm Bruslé and Quignard, 2001
16 Length at sexual maturity 9.3 [Female] 9.3 cm Fishbase, 2006
16 Length at sexual maturity 5.5 5.5 cm Rosecchi, 2001
16 Length at sexual maturity Female were mature at 9.29 in Stour 9.29 cm Mann, 1980
16 Length at sexual maturity Mean of 11.04, range 8.5-13.6 for females studied 11.05 cm Banbura and Koszalinski, 1991
15 Age at sexual maturity 2-3 [Both sex] 2.5 year Kennedy and Fitzmaurice, 1972
15 Age at sexual maturity 3 3.0 year Bruslé and Quignard, 2001
15 Age at sexual maturity 2 [Sometimes 1, rarely 3-4, Box sex] 3.5 year Rosechhi and Kestemont, 2001
15 Age at sexual maturity 2-3 [Female] 2.5 year Fishbase, 2006
15 Age at sexual maturity 1 [Bot sex] 1.0 year Rosecchi, 2001
15 Age at sexual maturity 1-2 [12-24 months, age at maturation] 1.5 year Vila-Gispert and Moreno-Amich, 2002
15 Age at sexual maturity 2-3 [Not specified] 2.5 year Environment agency, 1996
15 Age at sexual maturity Nearly all fish were mature at the end of their third year, and approximatively 89% from females and 74% Stour females spawnerd after two years 3.0 year Mann, 1980
15 Age at sexual maturity Celle-ci peut-être atteinte dès le second été (1+) chez les groupes qui mesurent 90 à 100 mm de longueur. […] 50% seulement des femelles de goujons 1+ sont aptes à se reproduire. Toutes les femelles 2+ sont aptes à la reproduction 1.0 year Brunet and Hoestlandt, 1972

Male (100%)


Trait id Trait Primary Data Secondary Data References
30 Male sexual dimorphism During the spawning season, male bears nuptial tubercules Present Spillmann, 1961
30 Male sexual dimorphism At spawning time, the heads of ripe male are thickly sprinkled with small nuptial tubercles like those of the dace. Nuptial tubercules also occur on the pectoral fins Present Kennedy and Fitzmaurice, 1972
30 Male sexual dimorphism Bears nuptial tubercules Present Bruslé and Quignard, 2001
30 Male sexual dimorphism During the spawning season, male bears nuptial tubercules on head and opercules Present Rinchard, 1996
30 Male sexual dimorphism In males, tubercles are much more intensively developped than in females. In larger individuals, the whole head, trunk, and all the fins are covered by tubercules [only head for smallest] Absent Witkowski and Rogowska, 1991
30 Male sexual dimorphism Male bears nuptial tubercles on head Present Billard, 1997
30 Male sexual dimorphism During the spawning season, male bears nuptial tubercules Present Rossechi and Kestemont, 2001
31 Onset of spermatogenesis From December increase regularly ['December'] Kestemond, 1989
31 Onset of spermatogenesis Developed gradually through the winter ['February', 'March', 'January'] Mann, 1980
33 Maximum GSI value 2.3 [June, prior to spawning] 2.3 percent Kestemond, 1989
33 Maximum GSI value 2.3 [During the breeding season] 2.3 percent Rosecchi, 2001
33 Maximum GSI value Huge variations between individuals: mostly 1-3% in May 2.0 percent Mann, 1980
32 Main spermatogenesis activity May and mid-June ['May', 'June'] Kestemond, 1989
32 Main spermatogenesis activity Increased rapidly in size just prior to spawning in late May and early June ['May', 'June'] Mann, 1980
35 Resting period 0.9 [September to October] 0.9 months Kestemond, 1989
34 Spermatogenesis duration During autumn and finish at the end of spring No data Kestemond, 1989
28 Length at sexual maturity 6.9 is the smallest male 6.9 cm Kennedy and Fitzmaurice, 1972
28 Length at sexual maturity 9.70-10.50 [Male] 10.1 cm Fishbase, 2006
28 Length at sexual maturity Mean size of mature males 10.46 cm in Frome and 9.66 cm in Stour 10.46 cm Mann, 1980

Spawning conditions (87%)


Trait id Trait Primary Data Secondary Data References
47 Mating system Si nos observations sont confirméesnle mode de reproduction du goujon pourrait être considéré comme polygame, chaque femelle étant suceptible de se reproduire avec plusieurs mâles successivement, voire même simultanément, et chaque mâle pouvant se reproduire successivement avec des femelles différentes No category Poncin, 1997
46 Nycthemeral period of oviposition Night Night Spillmann, 1961
46 Nycthemeral period of oviposition Le goujon pond normalement au début de la nuit Night Brunet and Hoestlandt, 1972
50 Parental care Nonguarder No care Fishbase, 2006
50 Parental care Non-guarder No care Mann, 1996
50 Parental care No parental protection of zygotes, embryo and larvae No category Vila-Gispert and Moreno-Amich, 2002
44 Spawning substrate Plants or gravels Ambiguous Spillmann, 1961
44 Spawning substrate Pebbles, weeds, tree roots and sodden leaves Ambiguous Kennedy and Fitzmaurice, 1972
44 Spawning substrate Psmanophile: sand, pebbles and small pebbles but sometimes aquatic plants Ambiguous Bruslé and Quignard, 2001
44 Spawning substrate Psmanophil: sand or gravels Ambiguous Rinchard, 1996
44 Spawning substrate Gravels and plants Ambiguous Billard, 1997
44 Spawning substrate Clean gravel and vegetation in flowing water Ambiguous Fishbase, 2006
44 Spawning substrate Eggs are laid on sand or fine roots associated with sand, washed by running water Ambiguous Mann, 1996
44 Spawning substrate Lithophil Lithophils Kennedy, 1969
44 Spawning substrate Gravel, typically 10-20 mm Lithophils Environment agency, 1996
44 Spawning substrate Psammophil Psammophils Wolter and Vilcinskas, 1997
44 Spawning substrate Psammophils Psammophils Balon, 1975
44 Spawning substrate Psammophil Psammophils Cattanéo, 2001
44 Spawning substrate Si nos résultats préliminaires sur le goujon tendaient à se confirmer, il deviendrait intéressant de discuter le caractère psammophile de la reproduction de cette espèce Psammophils Poncin, 1997
44 Spawning substrate Sur un fond de gravier ou de cailloutis […] Accessoirement les œufs peuvent être pondus, partiellement au moins, sur les végétaux immergés des rives du cours d'eau. Il est probable que les œufs qui tombent sur un fond vaseux sont perdus, car en aquarium, on constate que des oeufs reposant sur le fond (verre ou sable) ne se développent pas No category Brunet and Hoestlandt, 1972
45 Spawning site preparation No No category Bruslé and Quignard, 2001
45 Spawning site preparation Open water/susbtratum egg scatterers Open water/substratum scatter Fishbase, 2006
45 Spawning site preparation No, eggs are laid on substrate [Open substratum spawners] Ambiguous Mann, 1996
45 Spawning site preparation Zygotes are placed in a special habitat (e.g. scattered on vegetation, or buried in gravel) Susbtrate chooser Vila-Gispert and Moreno-Amich, 2002
45 Spawning site preparation Open substratum spawner Open water/substratum scatter Balon, 1975
45 Spawning site preparation Ce comportement a pour effet de libérer les œufs dans la colonne d'eau No category Poncin, 1997
41 Spawning temperature 14-17 [But sometimes 12°C] 15.5 °C Kennedy and Fitzmaurice, 1972
41 Spawning temperature Ovulation requires a temperature of 16-17 [Optimum 20°C] 16.5 °C Bruslé and Quignard, 2001
41 Spawning temperature 14-17 15.5 °C Rinchard, 1996
41 Spawning temperature 16-17 is the minimum required 16.5 °C Kestemont, 1987
41 Spawning temperature Above about 15°C 15.0 °C Rosechhi and Kestemont, 2001
41 Spawning temperature 12-17 14.5 °C Mann, 1996
41 Spawning temperature 17 and more 17.0 °C Environment agency, 1996
41 Spawning temperature Spawning occured at daily mean temperature of 13.6 to 16.5°C. Based on all studies, it could be concluded that spawning occurs most frequently in May and June at water temperatures between 15 and 18°C 13.6 °C Penaz and Prokes, 1978
41 Spawning temperature 12-17 14.5 °C Kamler and Wolnicki, 2006
41 Spawning temperature La température de l'eau était comprise entre 22 et 24°C et la photopériode fixée à 16L/8N 22.0 °C Poncin, 1997
41 Spawning temperature La date des premières pontes annuelles est essentiellement dépendante de la température de l'eau (+ 16 à + 17°C), 16.0 °C Brunet and Hoestlandt, 1972
40 Spawning period duration 4-8 6.0 weeks Bruslé and Quignard, 2001
40 Spawning period duration 4-8 [A female spawns about 4 times, with an interval of 1-2 weeks between each] 6.0 weeks Kestemont, 1987
40 Spawning period duration 4-16 [1.00-4.00 month, length of breeding season] 10.0 weeks Vila-Gispert and Moreno-Amich, 2002
40 Spawning period duration In 1976: 11-13 May, at a mean watre temperature of 15.4 to 16.4°C, 17-19 May, at 15.1-16.4°C, 25-27 May, at 14.5-16.5°C and 10-12 June at 13.6 to 16.0°C 12.0 weeks Penaz and Prokes, 1978
42 Spawning water type Rivulet of about 30 cm wide, canal of 2 m wide Stagnant water Kennedy and Fitzmaurice, 1972
42 Spawning water type Slow-flowing current : 10 to 80 cm/s Flowing or turbulent water Bruslé and Quignard, 2001
42 Spawning water type Water with some current Flowing or turbulent water Spillmann, 1961
42 Spawning water type Current velocity: 2-80 cm/s Flowing or turbulent water Mann, 1996
42 Spawning water type Dans une eau faiblement courante Flowing or turbulent water Brunet and Hoestlandt, 1972
43 Spawning depth 5 to 8 cm deep and 40-50 cm deep 45.0 m Kennedy and Fitzmaurice, 1972
43 Spawning depth Shallow waters No data Bruslé and Quignard, 2001
36 Spawning migration distance Limited home range No data Environment agency, 1996
39 Spawning season May-June ['May', 'June'] Billard, 1997
39 Spawning season Mid-April until End of August, with a peak in May-June ['April', 'August', 'May', 'June'] Rinchard, 1996
39 Spawning season April-May to June-July and Mid-August ['April', 'May', 'August', 'June', 'July'] Bruslé and Quignard, 2001
39 Spawning season April to June [France] ['April', 'June'] Kennedy and Fitzmaurice, 1972
39 Spawning season May-June ['May', 'June'] Spillmann, 1961
39 Spawning season April to July [Peak in May-june] ['April', 'May', 'July'] Rosechhi and Kestemont, 2001
39 Spawning season May-July ['May', 'July', 'June'] Mann, 1996
39 Spawning season April-July ['April', 'May', 'July', 'June'] Rosecchi, 2001
39 Spawning season April-August ['April', 'May', 'August', 'June', 'July'] Environment agency, 1996
39 Spawning season Spawn in France between April and May. In the Don River, spawning was observed at the turn of April and May. In the waters of Ireland, spawn from lata May till late June ['April', 'May', 'June'] Penaz and Prokes, 1978
39 Spawning season (April) May-June ['April', 'May', 'June'] Kamler and Wolnicki, 2006
39 Spawning season May-June ['May', 'June'] Cattanéo, 2001
39 Spawning season In the Trent, the main hatching period for gudgeon was between late May and mid-June, with smaller numbers of fish from subsequent cohorts appearing at intervals therefater ['May', 'June'] Nunn, 2007
39 Spawning season Pond au printemps et en été […] elle variera donc selon la latitude, l'altitude et les différences annuelles des températures printanières […] C'est ainsi que dans les Dombes, la ponte peut commencer dès la seconde quinzaine d'avril alors que dans la Montagne Noire il faut attendre la seconde quinzaine de juin ou même le début de juillet. La période de ponte s'échève début juin dans les eaux relativement chaudes alors qu'elle peut durer jusqu'en septembre dans les eaux froides. Dans le Sud de l'Angleterre, la ponte peut débuter en fin avril (Tamise), mais en Irlande, elle ne commencera qu'en mai ou en juin No data Brunet and Hoestlandt, 1972
48 Spawning release Frationnal and multiple spawner Mutliple Chemillier, 1995
48 Spawning release Frationnal and multiple spawner Mutliple Rossechi and Kestemont, 2001
48 Spawning release Batch spawner [Spawns once a year in low productivity streams, but exhibits multiple spawinng within a season in high productivity environments] Ambiguous Fishbase, 2006
48 Spawning release By batches Mutliple Spillmann, 1961
48 Spawning release Ova were scattered over the bed of the rivulet singly, in two and three and occassionaly in clumps of 4 together No category Kennedy and Fitzmaurice, 1972
48 Spawning release 1000 to 5000 eggs are released each time, and 8-15 days between each spawning and four spawning duraing the season No category Bruslé and Quignard, 2001
48 Spawning release Multiple spawner Mutliple Rinchard, 1996
48 Spawning release Up to four batches Mutliple Kestemont, 1987
48 Spawning release Released by small batches Mutliple Billard, 1997
48 Spawning release Either single spawning per year of two to four spawnings per year Total Vila-Gispert and Moreno-Amich, 2002
48 Spawning release Typical multiple spawners Mutliple Fredrich, 2003
48 Spawning release Most females lay two or more batches of eggs each season Mutliple Mann, 1980
48 Spawning release Spawned repeatedly in four terms No category Penaz and Prokes, 1978
48 Spawning release Fractional Fractional Cattanéo, 2001
48 Spawning release Adopt multiple spawning strategies, with up to three batches of eggs produced by individual fish Mutliple Nunn, 2007
48 Spawning release Ponte multiple Mutliple Beelen, 1998/1999
48 Spawning release Pluralité de pontes d'une même femelle au cours d'une même saison No category Brunet and Hoestlandt, 1972
49 Parity Spawns once a year for several years No category Fishbase, 2006
49 Parity Both sex had low survival rates ad their reproductive life spans were rarely more than three years No category Mann, 1980
49 Parity Peut vivre de 5 à 7 années et se reproduire dès la seconde année No category Brunet and Hoestlandt, 1972