Smyly, W.J.P. (1955) On the biology of the stone loach Nemacheilus barbatula (L.). Journal of Animal Ecology, pp. 167-186
| Species | Development state | Trait | Primary Data | Secondary Data |
| Barbatula barbatula | Egg | Egg size after water-hardening | Diameter of 1.0 mm and a tough outer skin | 1.0 mm |
| Barbatula barbatula | Egg | Egg Buoyancy | The eggs from these fish were found scattered over the bottom of the pond, the majority singly | Demersal |
| Barbatula barbatula | Egg | Incubation time | Took 14-16 days at 12-16°C | 15.0 days |
| Barbatula barbatula | Egg | Temperature for incubation | Took 14-16 days at 12-16°C | 15.0 °C |
| Barbatula barbatula | Egg | Degree-days for incubation | Took 14-16 days at 12-16°C | 15.0 °C * day |
| Barbatula barbatula | Larvae | Initial larval size | The newly hatched fry measured 3.0 mm in length | 3.0 mm |
| Barbatula barbatula | Larvae | Full yolk-sac resorption | The yolk-sac is absorbed within 5-7 days [4.0-6.5 mm in size] | 6.0 °C * day |
| Barbatula barbatula | Female | Female sexual dimorphism | In many fish, the pectoral fin is relatively long and pointed in the male and round and short in the female. In most adult fish, but not all, the male pectoral fin was longer than that of the female; the distinction is therefore a useful but not infaillible means of determining sex from external examination. Immature fish could not be separate by this way. In the breeding season, no change of colour in either sex has been seen but small papillae, present on the pectoral fins of only male fish, have been found | Absent |
| Barbatula barbatula | Female | Absolute fecundity | Total number of eggs varied between 5000 and 6000 of which usually rather then half were ripe | 5000.0 thousand eggs |
| Barbatula barbatula | Female | Onset of oogenesis | Between September and February GSI lies between 5 and 10% | ['February', 'September'] |
| Barbatula barbatula | Female | Intensifying oogenesis activity | From March to June, with a few small fish excepted, this ratio lies on or above 10%, in a few instances reaching the high figures of 35% | ['March', 'April', 'May', 'June'] |
| Barbatula barbatula | Female | Maximum GSI value | From March to June, with a few small fish excepted, GSI lies on or above 10%, in a few instances reaching the high figures of 35%. From July to August, the ratio ranges between 3 and 20%, the higher values being largely late-spawners or non-spawners | 10.0 percent |
| Barbatula barbatula | Female | Resting period | From July to August, the GSI ranges between 3 and 20%, the higher values being largely late-spawners or non-spawners | 3.0 months |
| Barbatula barbatula | Male | Maximum GSI value | The weight of testus was seldom more than 2% of the body weight | 2.0 percent |
| Barbatula barbatula | Spawning conditions | Spawning season | In the Lake District, the fish spawns in May, though some breeding may take place in April and June, and a few ripe females have been found as late as July [Seasonal changes in the ratio gonad weight: body weight supports the conclusion that May is the month when most fish breed] | ['April', 'May', 'June', 'July'] |
| Barbatula barbatula | Spawning conditions | Spawning water type | [In three successive years, fish in aquarium, in which the water was still, did not spawn; but a circular concrete ponds out of doors supplied with water from jets set at an angle so that the water in the pond was continually moving | Stagnant water |
| Barbatula barbatula | Spawning conditions | Spawning substrate | The eggs are not laid in holes but on stones and plants | Phytophils |
| Barbatula barbatula | Spawning conditions | Spawning site preparation | The eggs are not laid in holes but on stones and plants | No category |
| Barbatula barbatula | Spawning conditions | Nycthemeral period of oviposition | It may be assumed with some certainty that spawning takes place in darkness either by night or on dark days | Day |
| Barbatula barbatula | Spawning conditions | Mating system | In the spawning act males and females come to lie on their sides belly to belly, so that reproductive openings are close together and fertilization is made more certain | No category |