No detailed citation.
| Species | Development state | Trait | Primary Data | Secondary Data |
| Leuciscus cephalus | Female | Maximum GSI value | 9-10% [In Mid-May] | 9.5 percent |
| Leuciscus cephalus | Male | Onset of spermatogenesis | The redevelopment of gonads did not begin until September | ['September'] |
| Leuciscus cephalus | Male | Maximum GSI value | Most between 2.5-4, some almost 6% [June] | 3.25 percent |
| Leuciscus cephalus | Female | Age at sexual maturity | The majority of males are mature at age V though some are ripe as early as age III | No data |
| Leuciscus cephalus | Female | Resting period | June-September quiescent period, About 1% | 1.0 months |
| Leuciscus cephalus | Male | Main spermatogenesis activity | March, but increase quite regularly between September to May | ['March', 'May', 'September'] |
| Leuciscus cephalus | Female | Onset of oogenesis | The redevelopment of gonads did not begin until September | ['September'] |
| Leuciscus cephalus | Female | Intensifying oogenesis activity | Increase regularly from September to May, but most between March-April | ['April', 'March', 'May', 'September'] |
| Leuciscus cephalus | Male | Resting period | June-September quiescent period, About 1% | 1.0 months |
| Leuciscus cephalus | Female | Absolute fecundity | Lowest fecundity observed at 32155 for a female 41.5 cm long and 1253 g and highest 64658 for a female 46 cm long and 1660 g | 32155.0 thousand eggs |
| Leuciscus cephalus | Spawning conditions | Spawning season | Spawning occurred from the end of May into June | ['May', 'June'] |
| Leuciscus cephalus | Female | Weight at sexual maturity | Mean weight of 92 g [Age 4] and 174 [Age 5] | 92.0 kg |
| Leuciscus cephalus | Female | Length at sexual maturity | Size in different regions, for females varie between 19.4-22.2 [Age 4] and 24.1-25.1 [Age 5] and 15.2 and 20.8 for both sex at age 4 | 20.8 cm |
| Leuciscus cephalus | Male | Length at sexual maturity | Size in different regions, for males varie between 13.8-20.1 [Age 3] and 19.4-22 [Age 4] and 15.2 and 20.8 for both sex at age 4 | 16.95 cm |
| Leuciscus cephalus | Male | Weight at sexual maturity | Mean weight of 33 g [Age 3] and 92 g [Age 4] | 33.0 kg |
| Leuciscus cephalus | Female | Age at sexual maturity | Most females are mature at age VII and some at V or VI | No data |
| Leuciscus cephalus | Spawning conditions | Parity | Each fish appeared to spawn every year | No category |