Lobon-Cervia, J. and Fernadez-Delgado, C. (1984) On the biology of the barbel (Barbus barbus bocagei) in the Jaram River. Folia Zoologica, pp. 371-384
| Species | Development state | Trait | Primary Data | Secondary Data |
| Barbus barbus | Female | Age at sexual maturity | 50% of the female are mature in their sixth year, 40% in their seventh year and 100% in their > eighth | 50.0 year |
| Barbus barbus | Female | Length at sexual maturity | Females became mature at 18-20 | 19.0 cm |
| Barbus barbus | Female | Female sexual dimorphism | The difference in age at first spawing betwen males and females is 3-4 years | Absent |
| Barbus barbus | Female | Onset of oogenesis | New increase in the gonad weight is noted from September, especially in females | ['September'] |
| Barbus barbus | Female | Intensifying oogenesis activity | During spring (March-May) a rapid frowth of the gonads occurs until the next spawning | ['March', 'April', 'May', 'June'] |
| Barbus barbus | Female | Maximum GSI value | About 11%, up to 12 [In beginning of June] | 11.0 percent |
| Barbus barbus | Female | Resting period | There seems to be a two-month quiescent period (July-August) | 3.0 months |
| Barbus barbus | Male | Age at sexual maturity | 6% of males are mature in their second year, 92% in their third year and 100% in their > fourth | 6.0 years |
| Barbus barbus | Male | Length at sexual maturity | Males become mature at 7 | 7.0 cm |
| Barbus barbus | Male | Male sexual dimorphism | Male and female barbels reach approximatively the same value of Gi in the reproductive period, hich is very uncommon in cyprinid fishes | Absent |
| Barbus barbus | Male | Onset of spermatogenesis | A new increase in gonad weight, seems to be in February | ['February'] |
| Barbus barbus | Male | Main spermatogenesis activity | April-May | ['April', 'May'] |
| Barbus barbus | Male | Maximum GSI value | About 11 [Beginning of June] | 11.0 percent |
| Barbus barbus | Male | Resting period | No differences in term of GSI between September till January | 3.0 months |