Zaytsev, A.M. (1986) Embryonic-larval development of grayling, Thymallus thymallus, from Lake Ladoga. J. Ichthyol., pp. 123-32
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Egg | Oocyte diameter | Average 2.5, range 2.0-3.0 [Unfertilized eggs] | 2.5 mm |
| Thymallus thymallus | Egg | Egg size after water-hardening | 2-8.3.0 is the diameter of the egg itself, but the diameter of the swollen eggs with the perivetlline space is 3.25-4.66 mm ] | 3.96 mm |
| Thymallus thymallus | Egg | Incubation time | 16-21 [9.0-10.5°C] | 18.5 days |
| Thymallus thymallus | Egg | Degree-days for incubation | 160-170 [Under the usual incubation conditions (9.0-10.5°C), mass hatching of lake grayling occurred at 16-17 days] | 165.0 °C * day |
| Thymallus thymallus | Larvae | Initial larval size | 10.9-11.1 | 11.0 mm |
| Thymallus thymallus | Larvae | Larvae behaviour | Prolarvae concentrated in groups at the bottom in the corners on the trough, rarely mouving around | Demersal |
| Thymallus thymallus | Larvae | Reaction to light | They have negative phototaxis, and search for contact with the ground and the shelter | Photopositive |
| Thymallus thymallus | Larvae | Temperature during larval development | 9.0-10.5°C | 9.75 °C |
| Thymallus thymallus | Larvae | Full yolk-sac resorption | 130-150 [13-14 at 9.0-10.5°C] | 140.0 °C * day |
| Thymallus thymallus | Larvae | Onset of exogeneous feeding | 100-110 [10-11 days at 9.0-10.5°C] | 105.0 °C * day |