Kortet, R. and Taskinen, J. and Vainikka, A. and Ylönen, H. (2004) Breeding tubercles, Papillomatosis and dominance behaviour of male roach (Rutilus rutilus) during the spawning behaviour. Ethology, pp. 591-601
| Species | Development state | Trait | Primary Data | Secondary Data |
| Rutilus rutilus | Female | Length at sexual maturity | Females are often larger than males | No data |
| Rutilus rutilus | Male | Male sexual dimorphism | All the males studied, including the ones with no breeding tubercles, were sexually mature and ready to spawn | Absent |
| Rutilus rutilus | Spawning conditions | Spawning season | Spawning in spring | ['April', 'May', 'June'] |
| Rutilus rutilus | Spawning conditions | Spawning temperature | In the laboratory, the fish were maintained at 11°C in single-sex groups in circular 1500 l tanks to keep their spawning stage stable prior to ehavioural tests […] The water temperature in the stream tank was kept between 18 and 20°C | 11.0 °C |
| Rutilus rutilus | Spawning conditions | Spawning water type | Littoral waters, bays, creeks and small ponds, which are warm in spring | Stagnant water |
| Rutilus rutilus | Spawning conditions | Spawning depth | In relatively shallow waters | No data |
| Rutilus rutilus | Spawning conditions | Spawning substrate | Spawns on stones and vegetation | Lithophils |
| Rutilus rutilus | Spawning conditions | Mating system | In large groups | Promiscuity |
| Rutilus rutilus | Spawning conditions | Mating system | Roach breed using a lek-like mating system in which males can have bourgeois and/or parastic roles | No category |
| Rutilus rutilus | Spawning conditions | Parental care | Roach show no parental care for their offspring | No care |