Ovidio, M. and Parkinson, D. and Sonny, D. and Phippart, J-C. (2004) Spawning movements of European grayling Thymallus thymallus in the River Aisne (Belgium). Folia Zool., pp. 87-98
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning migration distance | Spawning migrations ranged from 70 to 4980 m | 70.0 km |
| Thymallus thymallus | Spawning conditions | Spawning migration period | Migrations started in March, under conditions of decreasing water level and increasing water temperature in a thermal range of 5 to 8°C | ['March'] |
| Thymallus thymallus | Spawning conditions | Spawning season | From 23 March to 11 April | ['March', 'April'] |
| Thymallus thymallus | Spawning conditions | Spawning period duration | About 3 weeks from 23 March to 11 April [Males stayed at the spawning grounds on average 12.2 ± 9.84 days and females 7.0 ± 7.6 days] | 12.2 weeks |
| Thymallus thymallus | Spawning conditions | Spawning temperature | 7-11 | 9.0 °C |