Witkowski, A. and Kowalewski, M. (1988) Migration and structure of spawning population of european grayling Thymallus thymallus (L.) in the Dunajec basin. Arch. Hydrobio., pp. 279-297
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Female | Age at sexual maturity | 2-3 [Most females were 3 years old] | 2.5 year |
| Thymallus thymallus | Female | Length at sexual maturity | Most frequently size of female 29.5-34.5, full range of 22.0-39.6 cm | 30.8 cm |
| Thymallus thymallus | Male | Age at sexual maturity | 1-2 for males, but age 2 dominates | 1.5 years |
| Thymallus thymallus | Male | Length at sexual maturity | Most frequently size of female 25.5-35.5, full range of 23.0-42.3 cm | 32.65 cm |
| Thymallus thymallus | Spawning conditions | Spawning migration period | When water temepratures is 4-6°C, which is most often at the end of March or at the beginning of April | ['March', 'April'] |
| Thymallus thymallus | Spawning conditions | Homing | In all fish marked in that year, as many as 18.7% came back to spawn to their home stream | Present |
| Thymallus thymallus | Spawning conditions | Spawning season | April | ['April'] |
| Thymallus thymallus | Spawning conditions | Spawning period duration | In very warm springs migrations and spawing takes 1.5-2 weeks long, if spring is cold last about a month | 1.75 weeks |
| Thymallus thymallus | Spawning conditions | Spawning water type | Spawn up rivers and streams, most often small clean tributaries | No category |