Haugen, T.O. and Vollestad, L.A. (2000) Population differences in early life-history in grayling. J. Evol. Biol., pp. 897-905
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Egg | Egg Buoyancy | Eggs are deposited a few centimetres below the gravel surface | Demersal |
| Thymallus thymallus | Egg | Temperature for incubation | 7.9-11.2 Reared conditions tested] | 9.55 °C |
| Thymallus thymallus | Egg | Degree-days for incubation | About 130-340 | 235.0 °C * day |
| Thymallus thymallus | Larvae | Larvae behaviour | The yolk_sac larvae remain buried until the yolk is exhausted | Demersal |
| Thymallus thymallus | Larvae | Full yolk-sac resorption | About 130 | 130.0 °C * day |
| Thymallus thymallus | Spawning conditions | Spawning migration period | Mature grayling from most lake-living populations migrate to streams and rivers after ice-break to spawn | No data |
| Thymallus thymallus | Spawning conditions | Homing | When mature, the grayling return to their natal streams with great precision to spawn | Present |
| Thymallus thymallus | Spawning conditions | Spawning season | Spring | ['April', 'May', 'June'] |