Nykänen, M. and Huusko, A. and Lahti, M. (2004) Movements and habitat preferences of adult grayling (Thymallus thymallus L.) from late winter to summer in a boreal river. Arch. Hydrobio., pp. 417-432
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning migration distance | Mean distances betwen winter site and spawning season site was 2744±1989 (range 130-5970 m ) | 2744.0 km |
| Thymallus thymallus | Spawning conditions | Spawning migration period | They leaft their overwintering sites during ice break-up and increasing spring fllod, at mean daily water temperature of 0.3-3.7°C, between 28 April to 2 May | ['January', 'February', 'March', 'April', 'May', 'June'] |
| Thymallus thymallus | Spawning conditions | Spawning season | The estimated spawning season lasted from 7 May to 1 June | ['May', 'June'] |
| Thymallus thymallus | Spawning conditions | Spawning period duration | About 2-3 [The estimated spawning season lasted from 7 May to 1 June] | 2.5 weeks |
| Thymallus thymallus | Spawning conditions | Spawning water type | Mean velocities of 40-70 | Flowing or turbulent water |
| Thymallus thymallus | Spawning conditions | Spawning depth | Seems to prefer water with water depth of 60-80 cm | 70.0 m |
| Thymallus thymallus | Spawning conditions | Nycthemeral period of oviposition | Spawning usually occurs in the afternoon or in the evening when water temperature is at his highest | Day |