Parkinson, D. and Philippart, J-C. and Baras, E. (1999) A preliminary investigation of spawning migrations of grayling in a small stream as detemrined by radio-tracking. Journal of fish biology, pp. 172-182
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning migration distance | Range from 230 to 4980 m | 230.0 km |
| Thymallus thymallus | Spawning conditions | Spawning migration period | All migrations started between 18 and 29 March | ['March'] |
| Thymallus thymallus | Spawning conditions | Homing | No certain, but in other areas grayligng ascened the same tributary year after year during their spawning runs | Present |
| Thymallus thymallus | Spawning conditions | Spawning period duration | About 2-3 [Males arrived on spawning grounds several days before females, and were detected on these grounds over much longer periods: 10-19 vs 2-3 days respecitively] | 2.5 weeks |
| Thymallus thymallus | Spawning conditions | Spawning temperature | 11 [Spawning behavior were observed on 28 March and lasted until 8 April] | 11.0 °C |
| Thymallus thymallus | Spawning conditions | Nycthemeral period of oviposition | Spawning events were noted only during the day, spawning can extend throughout the night in favourable thermal conditions | Day |