Pavlov, D.S. and Nezdolii, V.K. and Ostrovskii, M.P. and Fomin, V.K. (1998) Homing in the grayling Thymallus thymallus in the basin of the upper Volga. Journal of Ichthyology [J. Ichthyol.]., pp. 552-553
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning migration period | The spawning migration of grayling from the Volga to its tributaries usually occurs from late March to early May | ['March', 'May'] |
| Thymallus thymallus | Spawning conditions | Homing | High rates of homing in grayling in the basin of the Upper Volga | Present |