Meyer, L;. (2001) Spawning migration of grayling Thymallus thymallus (L., 1758) in a Nothern german lowland river. Arch. Hydrobio., pp. 99-117
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning migration distance | The maximum length of river sections used by the individual grayling ranged from 4.18-11.28 km | 7.73 km |
| Thymallus thymallus | Spawning conditions | Spawning migration period | Spawning run (12-25 March), a period of residency (26 March-1 April) and the dowstream migration (2-16 April) | ['March', 'April'] |
| Thymallus thymallus | Spawning conditions | Spawning water type | Upstream border of a shallow riffle, varied between 0.33-0.46 m/s | Flowing or turbulent water |
| Thymallus thymallus | Spawning conditions | Spawning depth | 0.55-1.15 m | 0.85 m |
| Thymallus thymallus | Spawning conditions | Spawning substrate | Lithophil: mainly coarse gravel, pebbles, cobbles and stones | Lithophils |
| Thymallus thymallus | Spawning conditions | Spawning site preparation | Dig shallow spawning pits (about 5 cm) compared to other salmonids | Susbtrate chooser |