Robards, M.D. and Quinn, T.P. (2002) The Migratory Timing of Adult Summer-Run Steelhead in the Columbia River over Six Decades of Environmental Change. Transactions of the American Fisheries Society [Trans. Am. Fish. Soc.], pp. 523-536
| Species | Development state | Trait | Primary Data | Secondary Data |
| Oncorhynchus mykiss | Spawning conditions | Spawning migration period | Enter freshwater from May to October in a sexually immature condition, remain in rivers all winter (often more than six months), and spawn the following spring | ['January', 'February', 'March', 'April', 'May', 'June', 'July', 'August', 'September', 'October'] |