Bengen, G. and Kugler, J. and Pequignot, J. . (1991) Study of the oocytes of Alosa alosa L. during spawning migration in the Garonne river. Cybium, pp. 229-238
| Species | Development state | Trait | Primary Data | Secondary Data |
| Alosa alosa | Female | Age at sexual maturity | 4-6 [Female] | 5.0 year |
| Alosa alosa | Male | Age at sexual maturity | 3-4 [Male] | 3.5 years |
| Alosa alosa | Spawning conditions | Spawning water type | Streams, water with current | Flowing or turbulent water |
| Alosa alosa | Spawning conditions | Spawning depth | 0.5-1.5 m | 1.0 m |
| Alosa alosa | Spawning conditions | Spawning substrate | Coarse gravel | Lithophils |