Nykänen, M. and Huusko, A. (2002) Suitability criteria for spawning habitat of riverine european grayling. Journal of fish biology, pp. 1351-1354
| Species | Development state | Trait | Primary Data | Secondary Data |
| Thymallus thymallus | Spawning conditions | Spawning water type | 40-70 cm/s | Flowing or turbulent water |
| Thymallus thymallus | Spawning conditions | Spawning depth | Depth < 40 cm were clearly avoided, prefereed 30-40 cm, nests were found at 38-106 cm deep | 35.0 m |
| Thymallus thymallus | Spawning conditions | Spawning substrate | Prefererred coarse gravel and fine pebbles [From sand, fine gravel, coarse gravel to fine cobble] | Lithophils |